Each InnoDB tablespace file (with a name matching *.ibd) is tagged with the file format used to create its table and indexes. The way to downgrade the tablespace is to re-create the table and its indexes. The easiest way to recreate a table and its indexes is to use the command:

ALTER TABLE t ROW_FORMAT=COMPACT;

on each table that you want to downgrade. The COMPACT row format uses the file format “Antelope”. It was introduced in MySQL 5.0.3.